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bahuvrIhau prakRtyA pUrva-padam

बहुव्रीहौ प्रकृत्या पूर्वॱपदम् ONPANINI 62001

In a /bahuvrIhi, the first half keeps its accent.

And the second member loses it.

So, an indra-kíller (/tatpuruSa compound) is someone who kills indra इन्द्र, but an índra-killer (/bahuvrIhi compound) is someone whose killer is indra इन्द्र, namely, someone who will be killed by indra इन्द्र.